Howard Eichenbaum

Howard Eichenbaum, Ph.D / Lab Chief

Howard Eichenbaum

The hippocampus plays a critical role in memory formation, but our understanding of just what the hippocampus does and how it performs its functions are still issues of considerable controversy. To enhance our knowledge about hippocampal function, we are pursuing a combination of neuropsychological studies of the nature of memory loss in animals with damage to the hippocampus and related cortical areas, and we are pursuing electrophysiological recording studies that seek to determine how information is represented by the hippocampus and associated cortical areas.

Howard Eichenbaum’s NeuroTree

Post Doctoral Associates


Zimbul Albo, Ph.D. – Post Doctoral Associate

Zimbul Albo

One of my primary research goals is to study the role of hippocampal ensembles and its related brain circuit in both spatial and non-spatial behaviors, with emphasis on naturalistic tasks such as social interaction and object-context recognition. I’m mostly excited about understanding the computational aspects of these functional ensembles and how this information passes during learning and memory to other related brain structures such as prefrontal cortex and thalamus. I think now is an exciting time to study population ensemble coding as calcium imaging, optogenetic and tetrode electrophysiology promise to reveal the mysteries of animal behavior.

David Sullivan

David Sullivan, Ph.D. – Post Doctoral Associate

David Sullivan

One day thirteen years ago, I decided not to cut class in order to attend a seminar on hippocampal neural network modeling, and I’ve been hooked on neuroscience ever since.  For my current project, I’m using genetic tools and miniaturized microscopes to examine the activity of many hippocampal neurons over long time periods.

Graduate Students

Ryan Placer

Ryan Placer, B.A. – Graduate Student

Ryan Place

Memory is life, as we know it. From sub-cellular processes necessary for enhanced connection, to the systems responsible for lasting cognition, I’d like to better understand how this representation of life is acquired, maintained and recalled. I’m currently fond of assessing the hippocampal formation by employing a combination of in vivo and behavioral techniques. Both up and down stream processing can be cool too.


Dan Salz

Dan Salz, B.A. – Graduate Student

Dan Salz

My research is looking to further our understanding of how the brain processes temporal information in relation to memory.  Recent studies have described a new phenomenon in pyramidal cells of dorsal CA1 in the Hippocampus of rats we’ve called “time cells”.  These cells fire at a specific time during some memory task delays similar to how place cells fire at certain locations in space. My project aims to help identify how and in which brain structures “time cells” originate, and to gain hints at how this information percolates through and is processed by the hippocampal system.

Dan Sheehan

Dan Sheehan, M.A. – Graduate Student

Dan Sheehan

I tend to focus my efforts on all things awesome.  As my graduate career is picking up momentum, awesome things include the ventral hippocampus.

Nat Kinsky

Nat Kinsky – Graduate Student

Nathaniel Kinsky

I am interested in the interplay between cortical areas and the hippocampus during the processes of consolidation/reconsolidation, when short-term memory representations transform/integrate into long-term representations.  Through the use of electrophysiological and imaging techniques, I hope to investigate the role that cortical areas, primarily the medial prefrontal cortex, play in this transformation/integration.

Graduate Student jhbladon(at)

John Bladon – Graduate Student jhbladon(at)

John Bladon

My background is in patch-clamp electrophysiology, and I have a MA in Anatomy and Neurobiology. I am interested in many things memory, but I focus specifically on the differences between spatial and non-spatial memories. To do this, I use in-vivo electrophysiology and optogenetics to examine the interactions between the hippocampus and cortical regions as the rat plays a memory game. In my free time I snowboard, rock climb, brew beer, and I play tennis and soccer.

Will Mau

Will Mau – Graduate Student wmau(at)

Will Mau

During my undergraduate years, I worked in a lab that utilizes extracellular recordings and behavior in rodents to study learning and memory. Since then, I have been addicted to behavioral neuroscience. My current research will use calcium imaging to study the basis for the formation of hippocampal “time cells” and how the responses of these cells evolve over learning. From these studies, I hope to gain a better understanding of the role of the hippocampal formation in episodic memory encoding and retrieval. My hobbies outside of the lab include dabbling in various physical activities such as rock climbing, Ultimate Frisbee, and extreme coffee drinking.

Catherine Mikkelsen

Catherine Mikkelsen – Graduate Student

Catherine Mikkelsen

Lab Alumni

Research Associates:

  • R. Jonathan Robitsek
  • Magdalena Sauvage
  • Norbert Fortin
  • Joe Manns
  • Amy Griffin
  • Paul Lipton
  • Inah Lee
  • Josh Berke
  • Seth Ramus
  • Jill McGaughy
  • Aras Petrulis
  • Russel Phillips
  • Pablo Alvarez
  • Emma Wood
  • Paul Dudchenko
  • Heikki Tanila
  • Tim Otto
  • Brian Young
  • Dario Dieguez, jr.
  • Murat Okatan
  • Andrea Frank
  • Rapeechai Navawongse
  • Lara Rangel
  • Jon Rueckemann
  • Marco Brockmann
  • Zachary Beer
  • Anja Farovik

Graduate Sudents

  • Benjamin Kraus
  • Audrey DiMauro
  • Rob Komorowski
  • Loren Devito
  • Michael Prerau
  • Cullen Owens
  • Bob Ross
  • Ceren Ergorul
  • Iain Wilson
  • Megan Libbey
  • Geoff Schoenbaum
  • Jeff Dusek
  • Archana Sheth
  • Hindiael Belchior
  • Sam McKenzie
  • Robert Robinson II
  • Suhani Mehrotra
  • Nick Robinson