Howard Eichenbaum

Howard Eichenbaum, Ph.D / Lab Chief

Howard Eichenbaum

The hippocampus plays a critical role in memory formation, but our understanding of just what the hippocampus does and how it performs its functions are still issues of considerable controversy. To enhance our knowledge about hippocampal function, we are pursuing a combination of neuropsychological studies of the nature of memory loss in animals with damage to the hippocampus and related cortical areas, and we are pursuing electrophysiological recording studies that seek to determine how information is represented by the hippocampus and associated cortical areas.

Howard Eichenbaum’s NeuroTree

James Priestley - Research Technician jpriestl(at)

James Priestley – Research Technician

James Priestley

I’m a recent graduate of BU’s undergraduate program in neuroscience and work primarily with Nick Robinson, studying sequential neuronal activity in the rat hippocampus through in-vivo electrophysiological and optogenetic interrogation of the hippocampal-entorhinal circuit.

Victor Kuo


Post Doctoral Associates

Anja Farovik, Ph.D - Post Doctoral Researcher af(at)

Anja Farovik, Ph.D – Post Doctoral Researcher

Anja Farovik

My work is centered around understanding the neural network responsible for the formation and retrieval of contextual memory representations. Using electrophysiology techniques to record neuronal activity patterns, current work explores how cells in the prefrontal cortex represent the context in which events happen.

Chris Keene

Chris Keene, Ph.D – Post Doctoral Associate

Chris Keene

My current research focuses primarily on the cortico-hippocampal processing stream providing contextual or “where” information to the hippocampus. Using a combination of electrophysiology and lesion techniques, this research will examine how cortical regions such as the entorhinal and retrosplenial cortices interact with the hippocampus and medial temporal lobe during learning and memory.


Jon Rueckemann, Ph.D – Post Doctoral Researcher

Jon Rueckemann

My research investigates how hippocampal afferents influence intrahippocampal processing, utilizing a combination of optogenetics, pharmacology, and extracellular recording in the behaving rat.

Marco Brockmann

Marco Brockmann, Ph.D – Post Doctoral Associate

Marco Brockmann

During my PhD in Germany, my research focused on prefrontal-hippocampal interactions in neonatal rats. At that stage, these regions already communicate with each other This is presumably a prerequisite for accomplishing higher cognitive functions in adulthood. Now, I have the chance to focus on these functions while rats perform behavioral tasks. My goal is to understand how neurons in the medial prefrontal cortex encode and retrieve information about the “what” and “where” features of memory.


Lara Rangel, Ph.D. – Post Doctoral Associate

Lara Rangel

One of the primary goals of my research is to better understand the different mechanisms through which cells in the dentate gyrus of the hippocampus can account for differences in spatial and non-spatial features of events, contexts, and their associations. I would like to understand how the representation of these differences at the single cell level ultimately contributes to discrimination abilities in behavior. Moreover, I am interested in understanding how input into the hippocampus is deferentially processed within the dentate gyrus, CA3, and CA1 subregions and how these structures work together in the service of learning. Lastly, I am interested in the rich oscillatory activity of the dentate gyrus, the modulation of these oscillations by learning, and what we can learn from single cell and local field potential relationships within and across structures.

David Sullivan

David Sullivan, Ph.D. – Post Doctoral Associate

David Sullivan

One day thirteen years ago, I decided not to cut class in order to attend a seminar on hippocampal neural network modeling, and I’ve been hooked on neuroscience ever since.  For my current project, I’m using genetic tools and miniaturized microscopes to examine the activity of many hippocampal neurons over long time periods.

Zimbul Albo


Zachery Beer

Zachery Beer, Ph.D. – Post Doctoral Associate

Zachery Beer

My current work focuses on establishing and optimizing the DREADDs technique (Designer Receptors Exclusively Activated by Designer Drugs). This technique offers a restricted spatio-temporal profile for the temporary inactivation of select brain regions. Of particular interest are target regions within the Medial Temporal Lobe and Prefrontal cortical areas. Future work will be focused on investigating the role of prefrontal and MTL sub-regions in the encoding, consolidation, and retrieval of complex object-context associations. It is hoped that this work will help elucidate the functionally distinct roles of these regions, which are vital to attention, learning, and memory.

Graduate Students

Ryan Placer

Ryan Placer, B.A. – Graduate Student

Ryan Place

Memory is life, as we know it. From sub-cellular processes necessary for enhanced connection, to the systems responsible for lasting cognition, I’d like to better understand how this representation of life is acquired, maintained and recalled. I’m currently fond of assessing the hippocampal formation by employing a combination of in vivo and behavioral techniques. Both up and down stream processing can be cool too.


Nicholas Robinson

Nicholas Robinson, B.A. – Graduate Student

Nick Robinson

Nicholas Robinson recently graduated from The University of Edinburgh’s Neuroscience undergraduate program where he developed a particular interest for the encoding, storage and retrieval of episodic memories. He has experience performing lesion studies and in vivo tetrode recordings from the hippocampus of behaving animals. He is also a keen tennis and squash player and enjoys hiking and rock climbing.

Dan Salz

Dan Salz, B.A. – Graduate Student

Dan Salz

My research is looking to further our understanding of how the brain processes temporal information in relation to memory.  Recent studies have described a new phenomenon in pyramidal cells of dorsal CA1 in the Hippocampus of rats we’ve called “time cells”.  These cells fire at a specific time during some memory task delays similar to how place cells fire at certain locations in space. My project aims to help identify how and in which brain structures “time cells” originate, and to gain hints at how this information percolates through and is processed by the hippocampal system.

Dan Sheehan

Dan Sheehan, M.A. – Graduate Student

Dan Sheehan

I tend to focus my efforts on all things awesome.  As my graduate career is picking up momentum, awesome things include the ventral hippocampus.

Nat Kinsky

Nat Kinsky – Graduate Student

Nathaniel Kinsky

I am interested in the interplay between cortical areas and the hippocampus during the processes of consolidation/reconsolidation, when short-term memory representations transform/integrate into long-term representations.  Through the use of electrophysiological and imaging techniques, I hope to investigate the role that cortical areas, primarily the medial prefrontal cortex, play in this transformation/integration.

Graduate Student jhbladon(at)

John Bladon – Graduate Student jhbladon(at)

John Bladon

My background is in patch-clamp electrophysiology, and I have a MA in Anatomy and Neurobiology. I am interested in many things memory, but I focus specifically on the differences between spatial and non-spatial memories. To do this, I use in-vivo electrophysiology and optogenetics to examine the interactions between the hippocampus and cortical regions as the rat plays a memory game. In my free time I snowboard, rock climb, brew beer, and I play tennis and soccer.

Will Mau

Will Mau – Graduate Student wmau(at)

Will Mau

During my undergraduate years, I worked in a lab that utilizes extracellular recordings and behavior in rodents to study learning and memory. Since then, I have been addicted to behavioral neuroscience. My current research will use calcium imaging to study the basis for the formation of hippocampal “time cells” and how the responses of these cells evolve over learning. From these studies, I hope to gain a better understanding of the role of the hippocampal formation in episodic memory encoding and retrieval. My hobbies outside of the lab include dabbling in various physical activities such as rock climbing, Ultimate Frisbee, and extreme coffee drinking.

Catherine Mikkelsen

Catherine Mikkelsen – Graduate Student

Catherine Mikkelsen

Lab Alumni

Research Associates:

  • R. Jonathan Robitsek
  • Magdalena Sauvage
  • Norbert Fortin
  • Joe Manns
  • Amy Griffin
  • Paul Lipton
  • Inah Lee
  • Josh Berke
  • Seth Ramus
  • Jill McGaughy
  • Aras Petrulis
  • Russel Phillips
  • Pablo Alvarez
  • Emma Wood
  • Paul Dudchenko
  • Heikki Tanila
  • Tim Otto
  • Brian Young
  • Dario Dieguez, jr.
  • Murat Okatan
  • Andrea Frank

Graduate Sudents

  • Benjamin Kraus
  • Audrey DiMauro
  • Rob Komorowski
  • Loren Devito
  • Michael Prerau
  • Cullen Owens
  • Bob Ross
  • Ceren Ergorul
  • Iain Wilson
  • Megan Libbey
  • Geoff Schoenbaum
  • Jeff Dusek
  • Archana Sheth
  • Hindiael Belchior
  • Sam McKenzie